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GENERAL BIOLOGY II (EVOLUTION, ECOLOGY & DIVERSITY)

COURSE DESCRIPTION


Course Code: BIO 102
Credit Units: 3

General Biology II is a continuation of the foundational biological sciences introduced in General Biology I. This course focuses on the principles of evolution, ecological relationships, and biological diversity. It examines how life originated, how species evolve over time through mechanisms of natural selection and genetic variation, and how organisms interact within ecosystems. Students are introduced to the patterns and processes that shape the diversity of life on Earth, the ecological principles that govern living systems, and the evolutionary relationships among organisms.

The course integrates classical evolutionary theory, modern genetics, environmental biology, population dynamics, community interactions, environmental impact, and conservation biology. It also explores the major domains of life—Bacteria, Archaea, and Eukarya—emphasizing their characteristics, classification, and ecological significance. Students will understand how biodiversity supports the stability of ecosystems, the threats facing global biodiversity, and strategies for conservation.


COURSE OBJECTIVES

At the end of this course, students should be able to:

  1. Describe the core principles and evidence supporting biological evolution.
  2. Explain mechanisms that drive evolutionary change, including natural selection, mutation, genetic drift, and gene flow.
  3. Analyze how populations adapt to their environments through evolutionary processes.
  4. Describe ecological principles such as energy flow, nutrient cycling, population dynamics, and ecosystem structure.
  5. Discuss ecological relationships between organisms, including competition, symbiosis, predation, and parasitism.
  6. Explain the hierarchical levels of biological organization from organisms to biosphere.
  7. Identify and classify the major groups of microorganisms, plants, fungi, protists, and animals.
  8. Explain the importance of biodiversity and the threats posed by human activities.
  9. Evaluate conservation strategies for protecting natural ecosystems and species.
  10. Apply ecological and evolutionary principles to real-world environmental challenges.

CHAPTER 1 

Introduction to Evolutionary Biology: Principles, Evidence & Mechanisms

1.0 Overview

Evolution is the central unifying concept of modern biology. It explains how the vast diversity of life on Earth emerged from common ancestors and how organisms change over generations. This chapter examines the foundations of evolutionary biology: historical perspectives, the evidence for evolution, mechanisms of evolutionary change, and the importance of evolution to biological sciences.


1.1 Historical Foundations of Evolutionary Thought

For centuries, humans attempted to explain the origin and diversity of life. Early theories were largely philosophical and not based on scientific evidence. One such belief was fixity of species—the idea that organisms were created in their present form and never changed. Another early misconception was spontaneous generation, which suggested that organisms could arise from non-living matter.

Lamarckism

Jean-Baptiste Lamarck (1744–1829) proposed one of the earliest theories of evolution. He believed:

  1. Organisms change toward perfection.
  2. Traits acquired during an organism’s lifetime could be passed to offspring.

For example, he argued that giraffes developed long necks because their ancestors stretched their necks to reach leaves, and the stretched trait was inherited. Although Lamarck’s mechanism was incorrect, he helped introduce the idea that species change over time.

Darwin and Wallace

Modern evolutionary biology was founded by Charles Darwin and Alfred Russel Wallace, who independently proposed the theory of natural selection.

Darwin’s 1859 book On the Origin of Species provided compelling evidence that species evolve by natural selection—a process where genetic variations that enhance survival and reproduction become more common in a population.


1.2 Evidence of Evolution

Multiple lines of scientific evidence support the theory of evolution:

1. Fossil Record

Fossils show progressive changes in organisms through geological time. Transitional fossils like Archaeopteryx demonstrate links between major groups (birds and reptiles).

2. Comparative Anatomy

Structural similarities among organisms reflect shared ancestry.

  • Homologous structures (e.g., human arm, bat wing) indicate common ancestry.
  • Analogous structures (e.g., wings of insects and birds) evolve independently.
  • Vestigial structures (e.g., human appendix) are remnants of structures that had important functions in ancestors.

3. Embryology

Embryos of vertebrates show striking similarities, suggesting a shared origin.

4. Molecular Biology

DNA and protein comparisons reveal how closely species are related.

5. Biogeography

Unique species found on islands (e.g., Galapagos finches) provide evidence of adaptive evolution.


1.3 Mechanisms of Evolution

Evolution occurs through changes in gene frequencies in a population over generations. The main mechanisms include:

1. Natural Selection

Individuals with favorable traits survive and reproduce more successfully.

Natural selection can occur in several forms:

  • Directional selection — one extreme is favored.
  • Stabilizing selection — average trait is favored.
  • Disruptive selection — both extremes are favored.

2. Mutation

Random changes in DNA introduce new genetic variations.

3. Genetic Drift

Random changes in allele frequencies, especially in small populations.

Types of drift:

  • Bottleneck effect — population drastically reduced by disaster.
  • Founder effect — small group colonizes new area.

4. Gene Flow

Movement of genes between populations through migration.

5. Sexual Selection

Traits that increase mating success become more common.


1.4 Speciation and Evolutionary Patterns

Speciation occurs when populations diverge into separate species.

Types of Speciation

  • Allopatric speciation: due to geographical isolation.
  • Sympatric speciation: occurs without physical barriers.

Patterns of Evolution

  • Adaptive radiation: rapid evolution into new forms (e.g., Darwin’s finches).
  • Convergent evolution: unrelated organisms develop similar traits.
  • Co-evolution: two species influence each other's evolution.

1.5 Importance of Evolution to Biology

Evolution explains:

  • antibiotic resistance
  • genetic diseases
  • crop improvement
  • biodiversity patterns
  • wildlife conservation

It forms the foundation of all biological sciences.


CHAPTER 2 

Ecological Principles: Ecosystems, Populations & Environmental Interactions

2.0 Overview

Ecology is the study of interactions between organisms and their environment. It examines how populations grow, how species interact, and how energy flows through ecosystems. This chapter focuses on ecological levels of organization, ecological relationships, ecosystem components, population dynamics, and energy flow.


2.1 Levels of Ecological Organization

Ecology is structured hierarchically:

  1. Organism — individual living entity.
  2. Population — group of organisms of the same species.
  3. Community — all interacting populations in an area.
  4. Ecosystem — community plus abiotic factors (soil, water, climate).
  5. Biome — large regions defined by climate (e.g., rainforest).
  6. Biosphere — all life on Earth.

2.2 Components of Ecosystems

Ecosystems consist of:

Biotic Components

  • Producers (plants)
  • Consumers (animals)
  • Decomposers (bacteria, fungi)

Abiotic Components

  • Temperature
  • Water
  • Light
  • Soil
  • pH
  • Climate

These components interact to sustain life.


2.3 Ecological Interactions

1. Competition

Occurs when organisms vie for the same resources such as food, mates, or territory.

2. Predation

One organism (predator) kills and eats another (prey). Predation influences population size and adaptation.

3. Symbiosis

Long-term relationships between species:

  • Mutualism (both benefit)
  • Commensalism (one benefits, other unaffected)
  • Parasitism (one benefits, other harmed)

4. Herbivory

Animals feed on plants; this drives plant defense mechanisms.


2.4 Energy Flow in Ecosystems

Energy flows through ecosystems in a one-way path:

Trophic Levels

  1. Primary producers
  2. Primary consumers
  3. Secondary consumers
  4. Tertiary consumers

Energy transfer is inefficient: about 10% is passed to the next level. This principle explains why top predators are few in number.


2.5 Biogeochemical Cycles

Major nutrient cycles:

1. Water Cycle — evaporation, condensation, precipitation.

2. Carbon Cycle — photosynthesis, respiration, combustion.

3. Nitrogen Cycle — nitrogen fixation, nitrification, denitrification.

4. Phosphorus Cycle — important for DNA, ATP, bones.

These cycles maintain ecosystem stability.


2.6 Population Ecology

Population Growth

Populations grow in patterns:

  • Exponential growth — rapid increase without limits.
  • Logistic growth — growth slows as population reaches carrying capacity.

Population Regulation

Factors include:

  • food supply
  • competition
  • predation
  • disease
  • climate

2.7 Human Impacts on Ecosystems

Humans influence ecosystems through:

  • deforestation
  • pollution
  • climate change
  • overfishing
  • habitat destruction

Conservation and sustainable practices are essential for ecosystem health.


CHAPTER 3 

Biological Diversity I: Microbial Life, Protists & Fungi

3.0 Overview

This chapter examines the first major branch of biological diversity: microorganisms. Microbes include bacteria, archaea, protists, and fungi. Though often microscopic, they form the foundation of life on Earth, contributing to decomposition, nutrient cycling, biotechnology, and health.


3.1 Domains of Life

All life belongs to three domains:

  1. Bacteria
  2. Archaea
  3. Eukarya

Bacteria and archaea are prokaryotes—simple cells lacking membrane-bound organelles. Eukarya includes protists, plants, fungi, and animals.


3.2 Bacteria

Characteristics

  • Prokaryotic
  • Cell walls with peptidoglycan
  • Asexual reproduction (binary fission)
  • Can form endospores
  • Extremely diverse

Shapes

  • Cocci (spherical)
  • Bacilli (rod-shaped)
  • Spirilla (spiral)

Nutrition Types

  • Autotrophs
  • Heterotrophs
  • Chemotrophs

Importance

Bacteria play vital roles:

  • nitrogen fixation
  • decomposition
  • biotechnology (insulin production)
  • gut microbiome

3.3 Archaea

Once grouped with bacteria, archaea are genetically distinct.

Characteristics

  • Extremophiles (live in harsh environments)
  • Unique membrane lipids
  • No peptidoglycan

Types include:

  • Methanogens
  • Halophiles
  • Thermophiles

Archaea are important for nutrient cycling and biotechnology.


3.4 Protists

Protists are eukaryotic organisms that are not plants, animals, or fungi.

Types

  • Protozoa (animal-like)
  • Algae (plant-like)
  • Slime molds (fungus-like)

Examples

  • Amoeba
  • Paramecium
  • Euglena
  • Plasmodium (malaria parasite)

Protists play roles in aquatic ecosystems and global oxygen production.


3.5 Fungi

Characteristics

  • Eukaryotic
  • Non-photosynthetic
  • Cell walls contain chitin
  • Decomposers
  • Reproduce sexually and asexually

Major Groups

  • Zygomycetes
  • Ascomycetes
  • Basidiomycetes
  • Deuteromycetes

Importance

  • decomposition
  • antibiotics (penicillin)
  • food production (yeast, mushrooms)
  • plant diseases

CHAPTER 4 

Biological Diversity II: Plants, Animals & Conservation Biology

4.0 Overview

This chapter finishes our exploration of biological diversity by examining the plant and animal kingdoms, their evolutionary adaptations, classification, and ecological significance. It concludes with an introduction to conservation biology.


4.1 Plant Diversity

Plants evolved from green algae and adapted to life on land.

Major Plant Groups

  1. Bryophytes (mosses)
  2. Pteridophytes (ferns)
  3. Gymnosperms (conifers)
  4. Angiosperms (flowering plants)

Plant Adaptations

  • vascular tissues
  • seeds
  • flowers
  • cuticles
  • stomata

Plants provide oxygen, food, habitat, and medicines.


4.2 Animal Diversity

Animals are multicellular eukaryotes that ingest food.

Major Animal Groups

  • Invertebrates (95% of animals)
  • Vertebrates (fish, amphibians, reptiles, birds, mammals)

Invertebrates

Includes:

  • sponges
  • cnidarians
  • mollusks
  • arthropods (insects)
  • echinoderms

Vertebrates

Possess a backbone.

Groups:

  • Fish
  • Amphibians
  • Reptiles
  • Birds
  • Mammals

Animals play key roles in food webs and ecosystem balance.


4.3 Biodiversity & Conservation

Importance of Biodiversity

  • ecosystem stability
  • food security
  • medicine
  • climate regulation

Threats to Biodiversity

  • habitat loss
  • climate change
  • pollution
  • invasive species
  • overexploitation

Conservation Strategies

  • protected areas
  • habitat restoration
  • captive breeding
  • environmental legislation
  • sustainable development

Conservation biology seeks to protect the planet’s biological heritage.


CHAPTER 1: FOUNDATIONS OF EVOLUTIONARY BIOLOGY (DETAILED LECTURE NOTES)


1.0 Introduction

Evolutionary Biology is the conceptual backbone of biological sciences. It explains how living organisms change over generations, how new species arise, and how the remarkable diversity on Earth came to be. Without evolution, biology would be a collection of unrelated facts. With evolution, all biological facts—from DNA sequences to ecosystems—become interconnected.

Evolution is defined as:
“A change in allele frequencies in a population over generations.”

These lecture notes delve into the foundations, evidence, mechanisms, patterns, and significance of evolution.


1.1 Development of Evolutionary Thought

1.1.1 Early Ideas About Life

Before the 18th century, biological thought was dominated by two ideas:

(a) Fixity of Species

The belief that species were created exactly as they appear today and are unchanging.

(b) Special Creation

It was believed that all organisms were created by a divine force in recent history.

(c) Spontaneous Generation

The belief that living things can arise from non-living matter (e.g., maggots arising from meat).
This idea persisted until scientists like Francesco Redi and Louis Pasteur disproved it.

These early beliefs lacked scientific foundation but shaped human understanding for centuries.


1.2 Lamarck’s Theory of Evolution

Jean-Baptiste Lamarck was the first scientist to propose a comprehensive theory of evolution (1809).

Lamarck’s Key Ideas

  1. Use and Disuse
    Organs used frequently become stronger; organs not used deteriorate.

  2. Inheritance of Acquired Characteristics
    Characteristics acquired during an organism’s lifetime could be passed to offspring.

Example:

Giraffes stretching their necks to reach leaves → longer necks inherited by offspring.

Evaluation

Although Lamarck’s mechanism was incorrect (genetic traits, not acquired traits, are inherited), his contribution is significant because:

  • He recognized environmental influence on organisms
  • He proposed that species are not fixed
  • He stimulated scientific interest in evolution

1.3 Darwin and Wallace: The Birth of Modern Evolution

1.3.1 Darwin’s Voyage on the HMS Beagle

In 1831, Charles Darwin embarked on a 5-year journey on the HMS Beagle as a naturalist.
He observed:

  • Fossils resembling living species
  • Unique species on isolated islands
  • Variation among individuals within the same species

His famous studies on the Galápagos finches were key to his insights.

1.3.2 Wallace’s Contribution

Alfred Russel Wallace independently developed a theory of natural selection.
He sent his manuscript to Darwin in 1858, prompting Darwin to publish On the Origin of Species (1859).


1.4 Evidence for Evolution

Multiple fields of biology provide overwhelming evidence.


1.4.1 Fossil Record

Fossils reveal the historical sequence of life. They show:

  • Transitional species (e.g., Archaeopteryx showing reptile-bird features)
  • Gradual changes in groups over time
  • Extinct species that once lived but no longer exist

Fossil Dating

  • Relative dating: age determined by rock layer
  • Radiometric dating: age determined using radioactive isotopes

1.4.2 Comparative Anatomy

(a) Homologous Structures

Structures with similar origin but different functions.
Example:

  • Human arm
  • Bat wing
  • Whale flipper
  • Cat forelimb

→ All share a common ancestor.

(b) Analogous Structures

Different ancestral origins but similar functions.
Example:

  • Bird wings vs. insect wings

→ Not evidence of shared ancestry, but of convergent evolution.

(c) Vestigial Structures

Structures with no apparent function.
Examples:

  • Human appendix
  • Pelvic bones in snakes
  • Wings of flightless birds

These are remnants of evolutionary history.


1.4.3 Embryological Evidence

Embryos of different animals show similarities that are not apparent in adults.

Example:
Human, fish, chicken, and frog embryos all have:

  • Gill slits
  • Tail structures

This suggests a common ancestry.


1.4.4 Molecular Biology

DNA comparisons provide some of the strongest evidence.

  • Humans and chimpanzees share 98–99% of DNA
  • Universal genetic code (DNA → RNA → Protein)
  • Similarities in proteins (e.g., hemoglobin) across species

This shows how closely organisms are related and how long ago they diverged.


1.4.5 Biogeography (Geographical Distribution of Species)

The distribution of organisms across continents provides evolutionary evidence.

  • Islands often have unique species (e.g., Galapagos, Hawaii)
  • Species on islands resemble species on nearest mainland

This supports the idea of common ancestry and migration.


1.4.6 Observed Evolution in Real Time

Examples

  • Bacteria developing antibiotic resistance
  • Insects evolving pesticide resistance
  • Rapid evolution in viruses (e.g., influenza)

Evolution is not just historical—it’s happening today.


1.5 Mechanisms of Evolution

Evolution occurs when genetic variations become more or less common in a population.

Major mechanisms include:


1.5.1 Natural Selection

Darwin’s central idea:
Organisms with advantageous traits survive and reproduce more successfully.

Requirements for Natural Selection

  1. Variation
  2. Heredity
  3. Differential survival and reproduction

Types of Natural Selection

(a) Directional Selection

Favors one extreme phenotype.
Example:
Peppered moths during the Industrial Revolution.

(b) Stabilizing Selection

Favors intermediate phenotype.
Example:
Human birth weight (too small/large = higher mortality).

(c) Disruptive Selection

Favors extreme phenotypes.
Example:
African seedcracker finches with small or large beaks.


1.5.2 Mutation

A mutation is a change in DNA sequence.
Mutations are the ultimate source of genetic variation.

Types:

  • Point mutations
  • Insertions/deletions
  • Chromosomal mutations

Mutations may be:

  • Beneficial
  • Neutral
  • Harmful

Only mutations in gametes (sex cells) are inheritable.


1.5.3 Genetic Drift

Random change in allele frequencies, especially in small populations.

(a) Bottleneck Effect

Population drastically reduced by disaster (e.g., flood, disease).
Surviving population may not represent original genetic diversity.

(b) Founder Effect

Small group colonizes a new habitat.
Example:
Amish populations showing high frequency of genetic disorders.


1.5.4 Gene Flow (Migration)

Movement of genes between populations through migration.
Increases genetic diversity and reduces differences between populations.


1.5.5 Sexual Selection

Traits that increase mating success become more common.

Examples:

  • Peacock’s tail
  • Bright coloration in male birds
  • Deer antlers

Sexual selection can produce extreme traits.


1.6 Speciation

Speciation is the formation of new species.

1.6.1 What Is a Species?

A species is a group of organisms that:

  • interbreed naturally
  • produce fertile offspring

1.6.2 Modes of Speciation

(a) Allopatric Speciation

Occurs when populations are geographically isolated.
Barriers include:

  • rivers
  • mountains
  • oceans
  • deserts

Isolation prevents gene flow → populations diverge → new species form.

(b) Sympatric Speciation

Occurs without physical barriers.

Mechanisms include:

  • polyploidy (common in plants)
  • ecological separation
  • sexual selection

1.7 Patterns of Evolution


1.7.1 Adaptive Radiation

Rapid evolution of many species from a common ancestor.

Examples:

  • Darwin’s finches
  • African cichlid fishes
  • Hawaiian honeycreepers

Adaptive radiation occurs when organisms exploit new ecological niches.


1.7.2 Convergent Evolution

Unrelated organisms develop similar traits due to similar environments.

Examples:

  • sharks (fish) and dolphins (mammals) → streamlined bodies
  • cactus (Americas) and euphorbia (Africa)

1.7.3 Co-evolution

Two species evolve in response to each other.

Examples:

  • flowers and pollinators
  • predators and prey
  • parasites and hosts

1.8 Human Evolution (Brief Introduction)

Humans share a common ancestor with apes.
Key evolutionary developments:

  • bipedalism
  • large brain
  • tool use
  • language

Fossil evidence includes Australopithecus, Homo habilis, Homo erectus, and Homo sapiens.


1.9 Importance of Evolution to Modern Biology

Evolution explains:

  • genetic diseases
  • antibiotic resistance
  • viral evolution
  • agricultural crop improvement
  • conservation biology
  • human origins
  • biodiversity patterns

Evolution is essential for modern medicine, ecology, and biotechnology.


MOLECULAR BASIS OF EVOLUTION

Understanding evolution at the molecular level provides the strongest evidence that all living organisms share a common ancestry. It also explains how small genetic changes accumulate to produce the vast biological diversity seen today.

1. DNA Structure and Mutation

DNA (deoxyribonucleic acid) is the hereditary material in almost all organisms. It carries genetic instructions for development, functioning, growth, and reproduction.

Mutations—changes in the DNA sequence—are the raw materials for evolution. They occur in several forms:

  1. Point mutations:A single nucleotide change.
    • Silent mutation: No effect on amino acid sequence.
    • Missense mutation: Changes one amino acid.
    • Nonsense mutation: Inserts a stop codon prematurely.
  2. Insertions or deletions (indels): Addition or removal of nucleotides.
  3. Gene duplication: Entire genes may be copied, giving evolution raw material to modify.
  4. Chromosomal mutations: Inversions, translocations, and polyploidy (extra sets of chromosomes in plants).

Most mutations are neutral; some are harmful, and a few are beneficial. Beneficial mutations are preserved by natural selection over time.

2. Genetic Variation

Variation within a population is essential for evolution to occur. Sources of genetic variation include:

  • Mutation (primary)
  • Meiosis (crossing over and independent assortment)
  • Random fertilization
  • Gene flow (migration introducing new alleles)

3. Gene Pool and Allele Frequencies

A population’s gene pool consists of all the alleles present in that population.

Evolution occurs when allele frequencies change from one generation to the next.

The Hardy–Weinberg equation helps determine whether a population is evolving:

p^2 + 2pq + q^2 = 1

where:

  •  = frequency of dominant allele
  •  = frequency of recessive allele
  • , ,  represent genotype frequencies.

A population is in Hardy–Weinberg equilibrium only if there is:

  • No mutation
  • No migration
  • Large population size
  • Random mating
  • No natural selection

These conditions almost never occur naturally, meaning evolution is always happening.


MECHANISMS OF EVOLUTION

Evolution operates through several powerful mechanisms that shape populations over time.

1. Natural Selection

Natural selection acts on phenotypes that enhance survival or reproduction. Key points:

  • Organisms produce more offspring than can survive.
  • Individuals vary in traits.
  • Variation is heritable.
  • Individuals with advantageous traits leave more offspring.

Types of natural selection:

  1. Directional selection: Favors one extreme phenotype.
    Example: Antibiotic resistance in bacteria.
  2. Stabilizing selection: Favors intermediate traits.
    Example: Human birth weight.
  3. Disruptive selection: Favors both extremes.
    Example: Birds with very large or very small beaks survive better than those with medium beaks.

2. Genetic Drift

Random changes in allele frequencies, especially in small populations.

Forms:

  • Bottleneck effect: Sudden reduction in population size (e.g., floods, disease).
  • Founder effect: A few individuals colonize a new area.

Drift reduces genetic variation.

3. Gene Flow

Movement of alleles between populations through migration.
It increases genetic diversity and prevents populations from becoming genetically distinct.

4. Mutation

The ultimate source of all new genetic variation. Although rare per gene, mutations accumulate across generations.

5. Non-random Mating

Includes inbreeding (mating between relatives) and assortative mating.
It increases homozygosity but does not directly change allele frequencies.


SPECIATION: THE FORMATION OF NEW SPECIES

Speciation is the process by which populations evolve to become distinct species.

1. Biological Species Concept

A species is a group of organisms that:

  • Interbreed in nature
  • Produce fertile offspring
  • Are reproductively isolated from other such groups

2. Reproductive Isolation

Pre-zygotic barriers (before fertilization):

  • Habitat isolation
  • Temporal isolation (different breeding times)
  • Behavioral isolation (courtship differences)
  • Mechanical isolation (anatomical incompatibility)
  • Gametic isolation (sperm and egg cannot fuse)

Post-zygotic barriers:

  • Hybrid inviability
  • Hybrid sterility (e.g., mule)
  • Hybrid breakdown

3. Modes of Speciation

A. Allopatric Speciation

Population becomes geographically separated.
Most common form.
Example: Darwin’s finches on different islands.

B. Sympatric Speciation

Occurs without geographical separation.
Common in plants via polyploidy.

C. Parapatric Speciation

Populations are adjacent but have limited contact; environmental differences drive divergence.


PATTERNS AND RATES OF EVOLUTION

Evolution can follow different tempos and patterns.

1. Gradualism

Evolution occurs slowly through small, accumulated changes.

2. Punctuated Equilibrium

Long periods of stability interrupted by bursts of rapid change.

3. Adaptive Radiation

One ancestral species evolves into many new species to exploit different ecological niches.
Examples:

  • Darwin’s finches
  • African cichlid fish
  • Mammals after dinosaur extinction

EVIDENCE FOR EVOLUTION

1. Fossil Record

Shows transitional forms (e.g., Archaeopteryx between reptiles and birds).

2. Comparative Anatomy

  • Homologous structures: Same structure, different function (human arm, bat wing).
  • Analogous structures: Different structure, same function (bat wing vs insect wing).
  • Vestigial organs: Reduced structures (human appendix).

3. Comparative Embryology

Closely related species show similar embryonic stages.

4. Molecular Evidence

DNA and protein sequence similarities reveal evolutionary relationships.

5. Biogeography

Distribution of species across continents supports evolutionary theory.
Example: Marsupials dominate Australia.


HUMAN EVOLUTION (SUMMARY)

Humans evolved from African primate ancestors.

Key genera:

  • Australopithecus (4–2 million years ago)
  • Homo habilis (first tool users)
  • Homo erectus (first to migrate out of Africa)
  • Homo neanderthalensis
  • Homo sapiens (modern humans, evolved ~300,000 years ago)

Important milestones:

  • Bipedalism
  • Brain expansion
  • Development of language
  • Cultural evolution

CHAPTER SUMMARY

Chapter 1 provided an in-depth review of evolution—its mechanisms, molecular foundations, historical development, and supporting evidence. Evolution remains one of the most powerful unifying theories in biology.


CHAPTER 2:

 ECOLOGICAL PRINCIPLES — POPULATIONS, COMMUNITIES & ECOSYSTEMS


2.0 Introduction

Ecology is the scientific study of how organisms interact with each other and with their environment. These interactions shape the distribution, abundance, behavior, and evolution of organisms. Ecology provides a framework for understanding natural systems, environmental challenges, and conservation strategies.

This chapter explores:

  • Levels of ecological organization
  • Ecosystem structure and function
  • Populations and communities
  • Energy flow and nutrient cycling
  • Biomes
  • Human impacts on ecological systems

Ecology is fundamental to solving global issues such as biodiversity loss, climate change, pollution, and resource depletion.


2.1 Levels of Ecological Organization

Ecology examines life at multiple hierarchical levels. Each level represents increasing complexity.


2.1.1 Organism Level

An organism is a single living individual—plant, animal, fungus, or microbe.
Organism-level ecology focuses on:

  • behavior
  • physiology
  • adaptations to the environment

For example, how desert plants conserve water or how birds migrate seasonally.


2.1.2 Population Level

A population is a group of individuals of the same species living in the same area at the same time.

Population ecology studies:

  • population size
  • density
  • distribution
  • growth rates
  • demographic patterns
  • reproductive strategies

Example: studying how lion populations change in the Serengeti.


2.1.3 Community Level

A community consists of all populations in a particular area.

Community ecology examines:

  • species interactions
  • food webs
  • competition
  • predation
  • symbiosis

Example: interactions between ants, acacia trees, and herbivores in grasslands.


2.1.4 Ecosystem Level

An ecosystem includes all living organisms (community) plus non-living components (abiotic factors).

Abiotic factors include:

  • sunlight
  • temperature
  • water
  • soil
  • nutrients

Ecosystem ecology studies:

  • energy flow
  • nutrient cycling
  • productivity

2.1.5 Biomes

A biome is a large ecological region characterized by particular climate patterns and major vegetation types.

Examples:

  • tropical rainforest
  • desert
  • savanna
  • tundra
  • temperate forest

Each biome hosts unique biodiversity and ecological processes.


2.1.6 Biosphere

The biosphere is the global ecosystem—all regions of Earth that support life.

It includes:

  • atmosphere
  • hydrosphere
  • lithosphere

The biosphere connects all living organisms into a single, dynamic system.


2.2 Components of an Ecosystem

Ecosystems have two major components: biotic and abiotic.


2.2.1 Biotic Components

Biotic components are the living parts of an ecosystem.

(a) Producers (Autotrophs)

Organisms that convert sunlight or chemical energy into food.

Examples:

  • green plants
  • algae
  • cyanobacteria

Producers form the base of all food chains.

(b) Consumers (Heterotrophs)

Organisms that eat other organisms.

Types:

  • Primary consumers (herbivores) — cows, rabbits, grasshoppers
  • Secondary consumers (carnivores) — snakes, foxes
  • Tertiary consumers (top predators) — lions, eagles, sharks

(c) Decomposers

Break down dead organic matter and release nutrients.

Examples:

  • bacteria
  • fungi
  • earthworms

Decomposers are essential for nutrient cycling.


2.2.2 Abiotic Components

Abiotic components determine the physical environment.

Examples include:

  • Light: controls photosynthesis and plant growth
  • Temperature: affects metabolic rate
  • Water: influences cell processes and organism distribution
  • Soil: provides minerals, nutrients, and stability
  • Wind: affects evaporation and pollination
  • pH: determines soil and water chemistry

Abiotic factors strongly influence which species can thrive in a given ecosystem.


2.3 Ecological Interactions

Species interact in various ways that shape community structure.


2.3.1 Competition

Competition occurs when organisms attempt to use the same resource.

Types of Competition

  • Intraspecific: within the same species
  • Interspecific: between different species

Competitive Exclusion Principle

Two species cannot occupy the same niche indefinitely.
One will outcompete the other.

Resource Partitioning

Species avoid competition by using different parts of the same resource.
Example: different bird species feeding at different heights on a tree.


2.3.2 Predation

Predation occurs when a predator kills and consumes prey.

Impacts of Predation

  • regulates prey populations
  • drives evolutionary adaptations
  • maintains ecological balance

Adaptations for Predation

Predators develop:

  • sharp teeth
  • claws
  • speed
  • venom

Prey develop:

  • camouflage
  • warning coloration
  • mimicry
  • defensive behavior

2.3.3 Symbiosis

Symbiosis is a long-term relationship between two species.

(a) Mutualism — both benefit

Examples:

  • bees and flowering plants
  • nitrogen-fixing bacteria and legumes
  • clownfish and sea anemones

(b) Commensalism — one benefits, other unaffected

Examples:

  • barnacles attaching to whales
  • birds nesting in trees

(c) Parasitism — one benefits, other is harmed

Examples:

  • tapeworms in humans
  • mistletoe on trees
  • mosquitoes feeding on blood

Parasites rarely kill their hosts immediately because they depend on them for survival.


2.3.4 Herbivory

Herbivory involves animals feeding on plants.

Examples:

  • cows grazing
  • insects chewing leaves

Plants respond with:

  • thorns
  • poison
  • tough leaves
  • chemical signals

Herbivory drives plant evolution and maintains diversity.


2.4 Energy Flow in Ecosystems

Energy flows in one direction, from the sun to producers to consumers.


2.4.1 Laws Governing Energy Flow

First Law of Thermodynamics

Energy cannot be created or destroyed.

Second Law of Thermodynamics

Energy transfer is inefficient; energy is lost as heat.


2.4.2 Trophic Levels

Trophic levels represent feeding positions:

  1. Producers
  2. Primary consumers
  3. Secondary consumers
  4. Tertiary consumers
  5. Decomposers

2.4.3 Food Chains and Food Webs

Food Chain

A linear pathway of energy flow.

Example:Grass → Grasshopper → Frog → Snake → Eagle

Food Web

Interconnected food chains.
More realistic than simple chains.


2.4.4 Ecological Pyramids

Three types:

(a) Pyramid of numbers

Shows number of organisms.

(b) Pyramid of biomass

Shows dry weight of organisms.

(c) Pyramid of energy (most accurate)

Shows energy content at each trophic level.
Only 10% of energy is passed to the next level.

This explains:

  • Why there are fewer top predators
  • Why herbivores outnumber carnivores
  • Why energy supply limits ecosystem size

2.5 Biogeochemical Cycles

Elements move through ecosystems in cycles.


2.5.1 Water Cycle

Processes:

  • evaporation
  • condensation
  • precipitation
  • infiltration
  • transpiration

Water is essential for life and climate regulation.


2.5.2 Carbon Cycle

Carbon moves through:

  • photosynthesis
  • respiration
  • decomposition
  • fossil fuel burning

Human activities have increased atmospheric CO₂, contributing to climate change.


2.5.3 Nitrogen Cycle

Nitrogen is required for:

  • DNA
  • proteins
  • chlorophyll

Major processes:

  • nitrogen fixation
  • nitrification
  • assimilation
  • ammonification
  • denitrification

Bacteria play the primary role.


2.5.4 Phosphorus Cycle

Phosphorus is found in:

  • bones
  • teeth
  • ATP

It cycles through rocks, soil, water, and organisms.
Phosphorus is often the limiting nutrient in freshwater ecosystems.


2.6 Population Ecology

Population ecology seeks to understand how and why populations change.


2.6.1 Population Characteristics

(a) Size — number of individuals

(b) Density — number per unit area

(c) Dispersion Patterns

  • Clumped — most common (e.g., herding mammals)
  • Uniform — seen in territorial species
  • Random — rare in nature

2.6.2 Population Growth

(a) Exponential Growth

Occurs under ideal conditions.
Results in J-shaped curve.
Example: bacteria.

(b) Logistic Growth

Growth slows as population approaches carrying capacity (K).
S-shaped curve.

Carrying Capacity

Maximum population size environment can support.


2.6.3 Factors Affecting Population Size

Density-Dependent

  • competition
  • predation
  • disease

Density-Independent

  • natural disasters
  • weather
  • pollution

2.7 Human Impacts on Ecosystems

Human activities significantly alter ecosystems.

Major Impacts

  1. Deforestation
  2. Pollution (air, water, soil)
  3. Climate change
  4. Overfishing
  5. Habitat destruction
  6. Invasive species
  7. Desertification

Consequences

  • species extinction
  • loss of biodiversity
  • reduced ecosystem services
  • altered climate patterns

Solutions

  • conservation strategies
  • sustainable development
  • renewable energy
  • reforestation
  • environmental policies

2.8 Importance of Ecology to Humans

Ecology helps us:

  • manage natural resources
  • understand environmental issues
  • improve agriculture
  • conserve biodiversity
  • predict effects of climate change

Ecology is vital for the survival of life on Earth. 



CHAPTER TWO: ECOLOGY



INTRODUCTION TO ECOLOGY

Ecology is the biological study of how organisms interact with each other and with their physical environment. It examines the relationships that link living organisms—plants, animals, fungi, and microorganisms—to the air, water, soil, and climate around them. The word “ecology” is derived from the Greek words oikos (house or environment) and logos (study), meaning the study of the home of organisms.

The fundamental idea in ecology is that no organism exists in isolation; all living things depend on one another and their environment for survival. Understanding ecology helps explain how life persists, how populations grow, why ecosystems change, and how human activities impact the natural world.

Ecology operates at multiple organizational levels:

  1. Organism
  2. Population
  3. Community
  4. Ecosystem
  5. Biome
  6. Biosphere

Each level provides a unique perspective on life and its interactions.


LEVELS OF ECOLOGICAL ORGANIZATION

1. Organism

An organism is an individual living thing. Its ecological study focuses on how it survives in its environment. This includes:

  • Behavioral adaptations
  • Physiological mechanisms
  • Structural features (morphology)

For example, a camel's humps store fat for energy during long periods without food, while its ability to conserve water helps it thrive in deserts.

2. Population

A population consists of organisms of the same species living in the same geographical area and capable of interbreeding. Populations have:

  • Size
  • Density
  • Growth rate
  • Age structure

Understanding population ecology helps scientists manage wildlife, protect endangered species, and control pests.

3. Community

A community is a collection of different populations interacting in the same environment. It includes all the organisms—plants, animals, and microorganisms—in a particular area.

Community interactions include:

  • Predation
  • Competition
  • Mutualism
  • Parasitism
  • Commensalism

For example, in a forest community, trees, birds, fungi, and insects coexist and interact in complex ways.

4. Ecosystem

An ecosystem includes all living organisms (biotic components) and their physical environment (abiotic components). Abiotic factors include:

  • Sunlight
  • Temperature
  • Water
  • Soil
  • Minerals
  • pH
  • Climate

Biotic and abiotic components interact through energy flow and nutrient cycling, forming a functional unit.

5. Biome

A biome is a large regional ecosystem characterized by distinct climate, vegetation, and animal life. Examples include:

  • Tropical rainforests
  • Deserts
  • Savannas
  • Temperate forests
  • Tundra

Biomes are shaped mainly by temperature and precipitation.

6. Biosphere

The biosphere is the global sum of all ecosystems. It includes all regions where life exists—land, water, and the atmosphere. It represents the “living skin” of the earth.


ABIOTIC FACTORS IN ECOLOGY

Abiotic (non-living) factors critically shape ecosystems and influence species distribution.

1. Sunlight

Sunlight is the primary source of energy for almost all ecosystems. It influences:

  • Photosynthesis
  • Temperature regulation
  • Behavioural patterns (e.g., migration, reproduction)

2. Temperature

Different organisms have specific temperature ranges for survival. For example:

  • Polar bears can withstand freezing conditions.
  • Thermophilic bacteria can survive at 90°C in hot springs.

Temperature also affects metabolic rates.

3. Water Availability

Water regulates metabolic activities and determines vegetation types.
Examples:

  • Cacti thrive in deserts with minimal water.
  • Rainforests support diverse species due to abundant rainfall.

4. Soil and Minerals

Soil quality affects plant growth.
Factors include:

  • pH value
  • Mineral content
  • Humus level
  • Soil texture (clay, sand, loam)

5. Climate and Weather Patterns

Long-term climate patterns determine biome distribution, while short-term weather affects daily organism activities.


ENERGY FLOW IN ECOSYSTEMS

Energy flows through ecosystems in one direction—from the sun to producers and then to consumers.

1. Trophic Levels

Ecosystem organisms are arranged into trophic (feeding) levels:

  1. Producers (Autotrophs): Plants, algae, some bacteria; convert sunlight to chemical energy via photosynthesis.
  2. Primary Consumers (Herbivores): Animals that eat plants.
  3. Secondary Consumers (Carnivores): Animals that eat herbivores.
  4. Tertiary Consumers: Top predators.
  5. Detritivores and Decomposers: Break down dead organic matter (fungi, bacteria).

2. Food Chains

A food chain illustrates the linear flow of energy. Example:

Grass → Grasshopper → Frog → Snake → Hawk

3. Food Webs

Food webs show interconnected food chains, providing a realistic view of ecosystem feeding relationships.

4. Ecological Pyramids

Ecological pyramids display the relative amounts of:

  • Energy
  • Biomass
  • Numbers of organisms

at different trophic levels.

Energy pyramids always have a wide base and narrow top because only 10% of energy is transferred from one level to the next; the rest is lost as heat.

This is known as the 10% Law of Energy Transfer.


BIOGEOCHEMICAL CYCLES

Nutrients are recycled in ecosystems through biogeochemical cycles.

1. Water Cycle

Processes:

  • Evaporation
  • Transpiration
  • Condensation
  • Precipitation
  • Runoff
  • Infiltration

The water cycle maintains freshwater availability.

2. Carbon Cycle

Carbon moves through:

  • Photosynthesis
  • Respiration
  • Decomposition
  • Fossil fuel combustion

Human activities like deforestation and burning fossil fuels increase atmospheric CO₂, contributing to climate change.

3. Nitrogen Cycle

Nitrogen-fixing bacteria convert atmospheric nitrogen into usable forms. Steps include:

  • Nitrogen fixation
  • Nitrification
  • Assimilation
  • Ammonification
  • Denitrification

Nitrogen is essential for proteins and nucleic acids.

4. Phosphorus Cycle

Unlike other cycles, phosphorus has no atmospheric component.
It cycles between rocks, soil, water, and organisms.


POPULATION ECOLOGY

Population ecology examines how populations grow and change.

1. Population Characteristics

  • Size (N)
  • Density (individuals per unit area)
  • Dispersion
    • Clumped (common in nature)
    • Uniform
    • Random
  • Age structure
    • Expansive (growing population)
    • Stationary
    • Constrictive (declining)

2. Population Growth Models

A. Exponential Growth

Occurs when resources are unlimited.
Produces a J-shaped curve.

Formula:

G = rN

B. Logistic Growth

Population growth slows as resources become limited.
Produces an S-shaped curve.

Formula:

G = rN \left(1 - \frac{N}{K}\right)

where:

  •  = carrying capacity
  •  = growth rate

3. Factors Affecting Population Size

Density-dependent factors

Increase with population density:

  • Competition
  • Disease
  • Predation

Density-independent factors

Affect populations regardless of density:

  • Floods
  • Drought
  • Wildfires

COMMUNITY ECOLOGY

Communities are shaped by interactions among organisms.

1. Competition

Occurs when organisms vie for the same resources.
Leads to:

  • Competitive exclusion
  • Resource partitioning

2. Predation

Involves a predator feeding on prey.
Prey develop adaptations like:

  • Camouflage
  • Mimicry
  • Speed
  • Defensive chemicals

3. Herbivory

Herbivores feed on plants.
Plants respond with spines, toxins, or rapid growth.

4. Symbiosis

Long-term interactions between species:

  • Mutualism (+/+)
    Example: Bees and flowers.

  • Commensalism (+/0)
    Example: Barnacles on whales.

  • Parasitism (+/–)
    Example: Tapeworms in humans.

5. Ecological Succession

Succession is a gradual change in community structure over time.

Primary succession

Occurs on bare surfaces (lava rocks, glaciers).

Secondary succession

Occurs after disturbances (fire, farming), where soil remains.

Climax community = stable final community (e.g., a mature forest).


ECOSYSTEM ECOLOGY AND BIOMES

1. Major World Biomes

A. Tropical Rainforest

  • High rainfall
  • High biodiversity
  • Dense vegetation

B. Savanna

  • Grasslands with scattered trees
  • Seasonal rainfall
  • Grazing animals dominate

C. Desert

  • Low rainfall
  • Plants adapted to drought
  • Animals active at night (nocturnal)

D. Temperate Forest

  • Distinct seasons
  • Deciduous trees

E. Grasslands

  • Dominated by grasses
  • Occasional wildfires
  • Rich soils

F. Tundra

  • Cold climate
  • Permafrost
  • Short growing seasons

ECOLOGY OF NATURAL RESOURCES

Natural resources include:

  • Water
  • Forests
  • Soil
  • Wildlife
  • Minerals

Ecology helps understand sustainable use by:

  • Conserving wildlife
  • Managing forests
  • Preventing soil erosion
  • Protecting watersheds

Human activities such as mining, pollution, overfishing, and deforestation threaten ecosystem stability.


HUMAN IMPACT ON THE BIOSPHERE

Human actions increasingly alter natural ecosystems.

1. Pollution

Types include:

  • Air pollution
  • Water pollution
  • Soil pollution

Consequences:

  • Acid rain
  • Eutrophication
  • Toxins entering the food chain

2. Deforestation

Leads to:

  • Loss of biodiversity
  • Climate change
  • Soil degradation
  • Disruption of water cycles

3. Climate Change

Caused mainly by increased greenhouse gases (CO₂, methane).
Results in:

  • Global warming
  • Sea-level rise
  • Extreme weather events
  • Habitat loss

4. Habitat Destruction

Major driver of species extinction.

5. Overexploitation

Overfishing, poaching, and unsustainable harvesting reduce species populations.


CONSERVATION ECOLOGY

Conservation biology aims to protect biodiversity and ecosystem health.

Key strategies:

  • Establishing protected areas
  • Reforestation
  • Wildlife breeding programs
  • Legislation (e.g., endangered species protection)
  • Environmental education
  • Sustainable resource management

The ultimate goal is to preserve life-support systems for future generations.


SUMMARY OF CHAPTER TWO

This chapter provided a detailed explanation of ecology, covering:

  • Levels of ecological organization
  • Abiotic and biotic factors
  • Energy flow and nutrient cycles
  • Population and community interactions
  • Ecosystem structure
  • Human impacts
  • Conservation strategies

Ecology emphasizes the interdependence of life and the need for responsible environmental stewardship.



CHAPTER THREE: BIOLOGICAL DIVERSITY (BIODIVERSITY)


INTRODUCTION TO BIOLOGICAL DIVERSITY

Biological diversity, commonly referred to as biodiversity, is the variety of life existing on Earth—encompassing genes, species, and ecosystems. It includes all plants, animals, microorganisms, and the ecological complexes in which they occur. Biodiversity is the product of over 3.5 billion years of evolution, shaped by natural selection, genetic variation, speciation, migration, and extinction.

The significance of biodiversity cannot be overstated. Life on Earth functions as an interconnected web where every organism, from microscopic bacteria to massive whales and giant trees, plays a role in sustaining natural systems. Biodiversity supports essential ecological processes such as nutrient cycling, pollination, decomposition, climate regulation, and soil formation.

This chapter provides a detailed examination of:

  • Levels of biodiversity
  • Classification of organisms
  • Origins of biological diversity
  • Major domains and kingdoms of life
  • Evolutionary relationships
  • Importance and threats to biodiversity
  • Conservation strategies

LEVELS OF BIODIVERSITY

Biodiversity exists at three interconnected levels:

1. Genetic Diversity

Genetic diversity refers to the variation of genes within a species. It includes:

  • Differences in DNA sequences between individuals
  • Variations in alleles
  • Genetic differences between populations

High genetic diversity increases a species’ ability to adapt to environmental changes, resist diseases, and survive disturbances. For example:

  • Wild populations of crops (e.g., rice, maize) have more genetic variation than cultivated varieties.
  • Human populations show diversity in skin color, blood groups, height, and susceptibility to diseases.

Loss of genetic diversity—often due to inbreeding, habitat fragmentation, or selective breeding—can make species more vulnerable to extinction.


2. Species Diversity

Species diversity refers to the variety of species within a region or ecosystem. It includes:

  • Species richness: Number of species
  • Species evenness: Relative abundance of species
  • Species dominance: Species with disproportionate ecological influence

Habitats such as tropical rainforests and coral reefs exhibit extremely high species diversity. For example:

  • The Amazon rainforest contains over 40,000 plant species and 3 million insect species.
  • Coral reefs host more species per unit area than any other marine environment.

Species diversity ensures ecosystem stability and resilience.


3. Ecosystem Diversity

Ecosystem diversity includes the variety of ecosystems in a region. It encompasses:

  • Forests
  • Grasslands
  • Wetlands
  • Deserts
  • Freshwater ecosystems
  • Marine habitats

Each ecosystem contains unique communities and ecological processes. Ecosystem diversity supports the global balance of energy flow, carbon storage, climate regulation, and nutrient cycling.


CLASSIFICATION AND TAXONOMY OF ORGANISMS

Biological classification (taxonomy) provides a systematic way to name, identify, and categorize organisms. Modern taxonomy uses evolutionary relationships (phylogeny) to arrange life forms.

1. Binomial Nomenclature

Developed by Carl Linnaeus, this system gives each species a two-part scientific name:

  • Genus name (capitalized)
  • Species name (lowercase)

Example: Homo sapiens, Panthera leo, Zea mays

Scientific names are universal and reduce confusion caused by local names.


2. Hierarchical Classification

Organisms are grouped into increasingly specific categories:

  1. Domain
  2. Kingdom
  3. Phylum
  4. Class
  5. Order
  6. Family
  7. Genus
  8. Species

A helpful mnemonic:
"Dear King Philip Came Over For Good Soup."

This hierarchy reflects evolutionary relationships, where organisms in the same group share common ancestry.


THE THREE DOMAINS OF LIFE

Advances in molecular biology divided all life into three major domains:

1. Domain Archaea

  • Ancient prokaryotes
  • Known for surviving in extreme environments: hot springs, high salinity, deep oceans
  • Unique cell membrane lipids
  • Include methanogens, halophiles, and thermophiles

Archaea are genetically distinct from bacteria.


2. Domain Bacteria

  • Prokaryotic organisms found in nearly every environment
  • Include the most abundant organisms on Earth
  • Some are pathogenic; others play essential ecological roles (e.g., nitrogen fixation, decomposition)

Examples: Escherichia coli, cyanobacteria.


3. Domain Eukarya

Organisms with true nuclei and membrane-bound organelles. This domain includes:

  • Protists
  • Fungi
  • Plants
  • Animals

Eukaryotes are more complex and diverse in structure and function.


THE FIVE MAJOR KINGDOMS OF LIFE

Traditionally, living organisms are grouped into five kingdoms:


1. Kingdom Monera (Prokaryotae)

  • Includes archaea and bacteria
  • Unicellular
  • No nucleus
  • Reproduce asexually
  • Some are autotrophic; others are heterotrophic

Examples: Lactobacillus, cyanobacteria.


2. Kingdom Protista

  • Eukaryotic organisms not classified as fungi, plants, or animals
  • Mostly unicellular
  • Include algae, protozoa, slime molds

Examples: Amoeba, Euglena, Paramecium.


3. Kingdom Fungi

  • Eukaryotic
  • Non-photosynthetic
  • Absorb nutrients from decaying matter
  • Cells contain chitin
  • Reproduce by spores

Examples: Mushrooms, yeasts, molds.

Fungi are essential decomposers in ecosystems.


4. Kingdom Plantae

  • Multicellular, eukaryotic, photosynthetic organisms
  • Contain chlorophyll
  • Make food through photosynthesis
  • Cell walls contain cellulose

Examples: Flowering plants, mosses, ferns, conifers.

Plants are primary producers in most ecosystems.


5. Kingdom Animalia

  • Multicellular eukaryotes
  • Heterotrophic
  • Lack cell walls
  • Specialized tissues and organs

Examples: Humans, insects, fish, mammals, reptiles.

Animals occupy diverse ecological niches as herbivores, carnivores, and decomposers.


ORIGIN AND EVOLUTION OF BIOLOGICAL DIVERSITY

Biodiversity emerged through billions of years of evolutionary processes.

1. Chemical and Biological Origins

Life likely began through:

  • Chemical evolution of organic molecules
  • Formation of protocells
  • Development of self-replicating molecules (RNA world hypothesis)

Earliest life was simple prokaryotes.


2. Evolution of Eukaryotes (Endosymbiotic Theory)

Proposed by Lynn Margulis.
States that:

  • Mitochondria originated from aerobic bacteria engulfed by larger cells
  • Chloroplasts originated from photosynthetic cyanobacteria

These bacteria lived symbiotically inside host cells and evolved into organelles.


3. Adaptive Radiation

Adaptive radiation is the rapid evolution of multiple species from a common ancestor, often following:

  • Mass extinction
  • Colonization of new habitats
  • Evolution of a key adaptation (e.g., wings in birds)

Examples:

  • Darwin’s finches
  • Mammals after dinosaur extinction
  • Cichlid fish in African lakes

4. Mass Extinctions

Earth has experienced five major mass extinctions, removing between 50–90% of species each time.
Examples:

  • Permian extinction (most severe)
  • Cretaceous extinction (dinosaurs)

After each extinction, surviving species diversify rapidly.


MAJOR GROUPS OF LIVING ORGANISMS

Biodiversity is often examined based on major multicellular groups:


1. Plants

Major plant groups include:

  • Bryophytes (mosses)
  • Pteridophytes (ferns)
  • Gymnosperms (conifers)
  • Angiosperms (flowering plants)

Plants are essential producers, contributing oxygen and food to ecosystems.


2. Animals

Animals are grouped into invertebrates and vertebrates.

Invertebrates

  • No backbone
  • Include insects, mollusks, worms, arachnids, echinoderms

Vertebrates

  • Possess backbone
  • Include fish, amphibians, reptiles, birds, mammals

Animals occupy every ecosystem on Earth.


3. Fungi

Essential decomposers that recycle nutrients.
Form mutualistic relationships:

  • Lichens (fungi + algae)
  • Mycorrhizae (fungi + plant roots)

4. Protists

A diverse group including:

  • Algae (photosynthetic)
  • Protozoa (animal-like)
  • Slime molds (fungi-like)

They serve as:

  • Producers in aquatic ecosystems
  • Decomposers
  • Parasites (e.g., Plasmodium, malaria)

5. Prokaryotes (Bacteria and Archaea)

Vital roles include:

  • Nitrogen fixation
  • Decomposition
  • Fermentation
  • Antibiotic production
  • Helping digestion (e.g., gut microbiota)

IMPORTANCE OF BIODIVERSITY

Biodiversity provides numerous ecological, economic, scientific, and cultural benefits.


1. Ecological Importance

  • Maintains ecosystem stability
  • Supports nutrient cycles
  • Enhances resilience to disturbances
  • Promotes energy flow
  • Provides habitats and ecological niches

2. Economic Importance

Biodiversity contributes to:

  • Agriculture
  • Fisheries
  • Tourism
  • Forestry
  • Pharmaceuticals (over 50% of drugs originate from natural sources)

3. Scientific and Educational Value

Studying biodiversity helps:

  • Understand evolution
  • Discover new species
  • Advance biotechnology

4. Cultural and Aesthetic Value

Many cultures derive identity, spirituality, and traditions from nature.


THREATS TO BIODIVERSITY

Human activities are accelerating biodiversity loss.

Key threats include:


1. Habitat Loss and Fragmentation

Caused by:

  • Deforestation
  • Urbanization
  • Agriculture
  • Mining

Leads to species displacement and extinction.


2. Pollution

Chemicals, plastics, oil spills, and pesticides harm wildlife.


3. Climate Change

Leads to:

  • Sea-level rise
  • Altered habitats
  • Temperature shifts
  • Changes in migration and reproduction patterns

Coral bleaching is a major consequence.


4. Overexploitation

Includes:

  • Overfishing
  • Poaching
  • Illegal wildlife trade
  • Unsustainable harvesting

5. Invasive Species

Non-native species outcompete native species.
Examples:

  • Water hyacinth in African wetlands
  • Cane toads in Australia

6. Disease

Emerging diseases threaten wildlife populations.


CONSERVATION OF BIODIVERSITY

Conservation involves protecting species, habitats, and ecosystems to maintain natural diversity.

1. In-Situ Conservation

Protecting species within their natural habitats.

Methods:

  • National parks
  • Wildlife reserves
  • Marine protected areas
  • Biosphere reserves

2. Ex-Situ Conservation

Protecting species outside their natural habitats.

Methods:

  • Zoos
  • Botanical gardens
  • Seed banks
  • Gene banks

3. Legislation

Examples:

  • Endangered Species Act
  • CITES (Convention on International Trade in Endangered Species)
  • National environmental laws

4. Sustainable Use of Resources

Includes:

  • Sustainable forestry
  • Organic farming
  • Responsible fishing
  • Renewable energy

5. Environmental Education

Public awareness drives conservation actions.


SUMMARY OF CHAPTER THREE

This chapter explored biodiversity in depth, covering:

  • Levels of biodiversity (genetic, species, ecosystem)
  • Classification and taxonomy of organisms
  • Evolutionary origins of diversity
  • Three domains and five kingdoms of life
  • Importance of biodiversity
  • Threats posed by human activities
  • Conservation strategies to protect natural systems

Biodiversity sustains life on Earth. Understanding its complexity and value is essential for environmental stewardship and for the survival of future generations.


CHAPTER 4


Chapter Overview

This chapter explores the full spectrum of biological diversity, from microorganisms to complex multicellular organisms. It explains how scientists classify life forms, the characteristics of the three domains of life, and the ecological importance of major groups such as plants, animals, fungi, protists, bacteria, and archaea. The chapter also highlights evolutionary relationships, phylogenetic trees, and how biodiversity underpins ecosystem functions.


LECTURE NOTES

4.1 Meaning of Biological Diversity (Biodiversity)

Definition

Biological diversity—commonly called biodiversity—refers to the full variety of life on Earth, including:

  1. Genetic diversity – variation in genes within a species
  2. Species diversity – variety of species in a region
  3. Ecosystem diversity – variation in habitats, communities & ecological processes
  4. Functional diversity – range of biological roles organisms perform

Life on Earth is estimated to include over 8.7 million species, though only about 2 million have been formally described.


4.2 Levels of Biodiversity

1. Genetic Diversity

  • Refers to the variation in DNA among individuals of a species.
  • Ensures population survival in changing environments.
  • Low genetic diversity increases risk of extinction (e.g., cheetahs, Irish potato famine).

2. Species Diversity

  • Refers to the number and abundance of species in a community.
  • Tropical rainforests and coral reefs are the most species-rich ecosystems.
  • High species diversity increases ecosystem stability and resilience.

3. Ecosystem Diversity

  • Includes forests, grasslands, deserts, wetlands, lakes, oceans, and more.
  • Each ecosystem hosts unique environmental conditions and species interactions.

4. Functional Diversity

  • The variety of ecological functions such as:
    • Photosynthesis
    • Decomposition
    • Nitrogen fixation
    • Pollination
  • Ecosystems with high functional diversity perform better under stress.

4.3 Importance of Biodiversity

Ecological Importance

  • Enhances ecosystem stability and productivity
  • Supports nutrient cycling, soil formation, pollination, seed dispersal
  • Provides natural pest control

Economic Importance

  • Food, medicine, fuelwood, fibers, industrial materials
  • Over 50% of pharmaceuticals originate from natural sources

Scientific Importance

  • Helps understand evolution and ecological relationships
  • Provides models for biotechnology and genetic engineering

Cultural & Aesthetic Value

  • Inspiration for art, religion, tourism, and recreation

4.4 Classification of Living Organisms

4.4.1 Why Do We Classify?

Classification helps in:

  • Organizing biological knowledge
  • Establishing evolutionary relationships
  • Identifying, naming, and describing organisms
  • Predicting characteristics shared by related organisms

4.4.2 The Taxonomic Hierarchy

Biological classification follows levels from broad to specific:

Domain → Kingdom → Phylum → Class → Order → Family → Genus → Species

Mnemonic: “Dear King Philip Came Over For Good Soup.”

Species

  • The fundamental unit of classification

  • Defined as organisms capable of interbreeding to produce fertile offspring

  • Scientific names use binomial nomenclature (Genus species):

    Example: Homo sapiens, Escherichia coli


4.5 Evolutionary Basis of Classification

Modern classification is based on phylogeny—evolutionary history and relationships.

Phylogenetic Trees

  • Diagram showing evolutionary relationships
  • Built using:
    • DNA sequences
    • Morphological features
    • Fossil evidence

Clades

A clade includes a common ancestor and all its descendants.
Cladistics helps identify monophyletic (true evolutionary) groups.


4.6 The Three Domains of Life

Life is organized into three major domains:

  1. Bacteria
  2. Archaea
  3. Eukarya

This system was introduced by Carl Woese based on molecular (ribosomal RNA) differences.


4.6.1 Domain Bacteria

Characteristics

  • Prokaryotic (no nucleus)
  • Single-celled
  • Cell wall contains peptidoglycan
  • Asexual reproduction (binary fission)
  • Can form spores and biofilms

Nutrition Types

  • Autotrophic (photosynthetic or chemosynthetic)
  • Heterotrophic (parasitic, saprophytic)

Ecological Importance

  • Decomposition
  • Nitrogen fixation (Rhizobium)
  • Pathogens (e.g., Mycobacterium tuberculosis)
  • Industrial use (yogurt, antibiotics, genetic engineering)

Examples

  • Cyanobacteria (blue-green algae)
  • Streptococcus, Staphylococcus
  • Escherichia coli

4.6.2 Domain Archaea

Unique Characteristics

  • Prokaryotic but genetically distinct from bacteria
  • Cell walls lack peptidoglycan
  • Membrane lipids with unique chemical structures
  • Live in extreme environments (extremophiles)

Major Groups

  • Thermophiles – hot springs
  • Halophiles – salty water
  • Methanogens – produce methane; live in anaerobic environments

Importance

  • Key role in carbon and nitrogen cycles
  • Used in biotechnology (enzymes for PCR such as Taq polymerase)

4.6.3 Domain Eukarya

Eukaryotic organisms have:

  • A nucleus
  • Membrane-bound organelles
  • Complex cell structures
  • Sexual reproduction (in many groups)

The domain includes four major kingdoms:

  1. Protista
  2. Fungi
  3. Plantae
  4. Animalia

4.7 Kingdom Protista

Protists are mostly unicellular eukaryotes, diverse in form and nutrition.

Groups

  1. Protozoa – animal-like, heterotrophic

    • Examples: Amoeba, Paramecium, Trypanosoma
  2. Algae – plant-like, photosynthetic

    • Examples: Chlamydomonas, Kelp, Diatoms
  3. Slime Molds – fungus-like

    • Decomposers

Importance

  • Produce substantial portions of global oxygen
  • Base of aquatic food chains
  • Some cause diseases:
    • Malaria (Plasmodium)
    • Sleeping sickness (Trypanosoma)

4.8 Kingdom Fungi

Characteristics

  • Eukaryotic
  • Heterotrophic absorbers
  • Cell wall made of chitin
  • Body consists of hyphae forming mycelium
  • Reproduce via spores

Types of Fungi

  • Molds – Rhizopus
  • Yeasts – Saccharomyces
  • Mushrooms – Agaricus
  • Parasitic fungi – cause plant diseases

Ecological Roles

  • Decomposition
  • Symbiosis (mycorrhizae)
  • Food and medicine production (penicillin, citric acid)

4.9 Kingdom Plantae

General Characteristics

  • Multicellular
  • Phototrophic (photosynthesis)
  • Cell wall made of cellulose
  • Chloroplasts with chlorophyll
  • Sexual and asexual reproduction

Major Plant Groups

1. Bryophytes (Non-vascular plants)

  • Mosses, liverworts
  • No true roots or vascular tissues
  • Require water for fertilization

2. Pteridophytes (Seedless vascular plants)

  • Ferns
  • Have vascular tissues (xylem & phloem)
  • Reproduce by spores

3. Gymnosperms

  • Naked seeds
  • Conifers, pine, cycad

4. Angiosperms

  • Flowering plants
  • Seeds enclosed in fruit
  • Most diverse plant group
  • Includes monocots and dicots

4.10 Kingdom Animalia

General Characteristics

  • Multicellular
  • Heterotrophic
  • Lack cell walls
  • Motile at some stage
  • Nervous and muscular systems (most groups)

Major Animal Groups

Invertebrates

  1. Porifera – Sponges
  2. Cnidaria – Jellyfish, Hydra
  3. Platyhelminthes – Flatworms
  4. Nematoda – Roundworms
  5. Mollusca – Snails, octopuses
  6. Annelida – Earthworms, leeches
  7. Arthropoda – Insects, crustaceans
  8. Echinodermata – Starfish

Vertebrates (Phylum Chordata)

  1. Fishes – bony and cartilaginous
  2. Amphibians – frogs
  3. Reptiles – snakes, lizards
  4. Birds – Aves
  5. Mammals – humans, primates

4.11 Microorganisms and Their Roles in Biodiversity

1. Viruses

  • Non-cellular infectious agents
  • Replicate only inside host cells
  • Examples: HIV, Influenza, Bacteriophages

2. Microbial Diversity in Ecosystems

  • Soil microbes support nutrient cycling
  • Gut microbiota assist digestion
  • Marine microbes form base of aquatic food webs

4.12 Threats to Biodiversity

Major Threats

  • Habitat loss
  • Climate change
  • Pollution
  • Overexploitation
  • Invasive species

Consequences

  • Species extinction
  • Reduced ecosystem services
  • Food insecurity
  • Increased disease outbreaks

4.13 Conservation of Biodiversity

Methods

  1. In-situ conservation

    • National parks
    • Wildlife reserves
    • Protected forests
  2. Ex-situ conservation

    • Zoos
    • Botanical gardens
    • Seed banks
  3. Legislation and Policies

    • Endangered Species Act
    • International treaties (e.g., CITES)
  4. Public education and awareness


4.14 Summary of Chapter 4

By the end of this chapter, students should understand:

  • What biodiversity is and why it matters
  • How organisms are classified based on evolutionary relationships
  • Characteristics of the three domains of life
  • Major kingdoms and representative organisms
  • The ecological roles of bacteria, archaea, protists, fungi, plants, and animals
  • Threats to biodiversity and conservation strategies.